Further reading:
http://support.microsoft.com/default.aspx?scid=kb;en-us;197132
Quote:
Infrastructure FSMO Role
When an object in one domain is referenced by another object in another domain, it represents the reference by the GUID, the SID (for references to security principals), and the DN of the object being referenced. The infrastructure FSMO role holder is the DC responsible for updating an object's SID and distinguished name in a cross-domain object reference.
NOTE: The Infrastructure Master (IM) role should be held by a domain controller that is not a Global Catalog server(GC). If the Infrastructure Master runs on a Global Catalog server it will stop updating object information because it does not contain any references to objects that it does not hold. This is because a Global Catalog server holds a partial replica of every object in the forest. As a result, cross-domain object references in that domain will not be updated and a warning to that effect will be logged on that DC's event log.
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